Born on March 27, 1971 and raised in Edmonton, Alberta, Nathan is the youngest son in a family of English teachers: parents Cookie and Bob Fillion, and older brother Jeff. Nathan attended the University of Alberta and planned to follow in his family's footsteps and was one semester short of graduation when he won the role of the charming Joey Buchanan on the ABC soap One Life to Live, and moved to New York City.

In addition to One Life to Live, Nathan also made appearances in several other ABC productions, including the made-for-TV movie Ordeal in the Arctic (1993) and as a series regular, Johnny Donnely, in Two Guys and a Girl (1998). In 2002, he won the lead role of war-hardened Captain Malcolm 'Mal' Reynolds on the short-lived but brilliant Fox Series, Firefly, a Joss Whedon creation. His work on Firefly also led to the role of preacher-gone-wrong and 'big bad', Caleb, in the last season of Whedon's Buffy The Vampire Slayer.

Aside from his success in the States, Nathan has appeared in a number of Canadian productions, including the television movie, Strange and Rich (1994), theatrical performances in Edmonton's Fringe Theatre Festivals,

here he was originally discovered, and he participated in Theatre sports as a member of Rapidfire Theatre, an Edmonton theater company. He also appeared in Die Nasty, a soap opera spoof-improv performed at a local Edmonton theater.

Nathan's recent work includes the filming of Serenity, the big-screen sequel to Firefly, which premiered Sept 30, 2005. He has also completed filming Slither, a horror film from James Gunn of Dawn of the Dead fame. Slither is set to be released on March 31st in U.S. theaters. Most recently, Nathan has finished filming a romantic comedy in L.A. titled Waitress, and stars opposite Kerri Russell. Waitress is the story of a pregnant, unhappily married waitress in the deep south who meets and falls into an unlikely relationship with a newcomer to her town as a last attempt at happiness. A release date has yet to be announced.

Nathan is 6'1" with brown hair and blue eyes, is currently single, and has been residing in Los Angeles since